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Here we aimed at uncovering the mechanisms underlying the implication of MBs in reversal learning. We reasoned that GABAergic feedback from the output region back to the input region of the MBs might be crucial to inhibit responses to previously reinforced odors during the reversal phase. We thus performed pharmacological experiments blocked GABAergic signaling in the MBs during olfactory reversal learning, by locally injecting antagonists of ionotropic or metabotropic GABA receptors, into the calyces or the vertical lobes. Our results show that GABAergic signaling is required for reversal learning but is dispensable for elemental learning. We thus provide a circuit-based explanation of the implication of MBs role in reversal learning.
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Then, we calculated for each social session, the percent change in response rate relative to the animal's average performance during alone sessions (individual Δ scores). One sample t-tests were used to determine whether these individual Δ scores significantly differed from 0, and paired t-tests to compare audience and coaction effects for each individual. Finally, because the results revealed variable Δs across groups, Pearson's correlations were used to test the idea of a link between mean Δs over the two social conditions and the degree of intimacy between observer and observee as objectivized by the index: I = years of shared housing/years of age (Table 1).
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Posts Figure 2. Response rate (mean ± sem) for the group (A) and for each individual (B). In (B), scores are illustrated for the alone condition (Al, dark gray bars), the two social conditions taken together (Soc, light gray bars), and for audience (o) and coaction (•) separately. ***p ≤ 0.001, *p ≤ 0.05.
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